Oaks of the World

Classifications List of
Back to
home page

(Hybrids A to F --- Hybrids G to L --- Hybrids S to Z)
x maccormickii Carruthers 1861 aliena x dentata
= dentata var. maccormickii (Carruth.) Skan 1899
Korea, N.E. China;
x maccormickoserrata
T.B.Lee 1961
aliena x dentata x serrata
x macdanielli Green & Hess 1996 robur X macrocarpa
pyramidal or rounded crown; more than 12 m tall; leaf 6-15 x 4-11 cm, glabrous and dark green above, minutely grey-green tomentose beneath; both apex and base variable; 5-7 lobes, with sinuses not reaching halfway to midrib; acorn 2 cm x 1.5, singly or paired, on a 3-6 cm long peduncle; cup enclosing 1/2 of nut, with silver grey velvetly tomentose scales; young shoot greenish grey, smooth, without corky ridges;
cultivar available : 'Heritage'
x macdonaldii Greene 1889

pacifica x lobata
= dumosa var. macdonaldii (Greene) Jepson 1923
= dumosa ssp macdonaldii (Greene) A.Camus 1935
island scrub oak ;
Santa Cruz, Santa Rosa and Santa Catalina Islands at < 600 m ;
2-15 m ; scaly and greyish bark; tomentose twigs; leaves deciduous 4-7 cm long, both ends rounded, with
2-4 shallow lobes, thick, stiff, tomentose beneath; acorn 2-3 cm, enclosed for 1/3 in hairy cup, maturing in 1 year;
-- absent on american mainland;
-- named after James Mac Donald who put up the money for the publication of "Illustration of West American Oaks" by Alb. Kellogg and Ed. Greene in 1889 ;
-- different from Q. lobata by shallower leaves sinus, smaller and sharper lobes; fixed hybrid;


x macnabiana Sudworth 1927 sinuata x stellata
= x mahonii E.J.Palmer 1937
Texas, Oklahoma, Arkansas ;
x major Nakai 1915 aliena x serrata
= glandulifera var. major Seem. ex Nakai 1915
x mannifera Lindl. 1840 infectoria ssp veneris x petraea ssp iberica
= x longifolia K.Koch 1849 nom. illeg.
= sessiliflora var. mannifera (Lindl.) Boiss. 1879
= robur var. mannifera (Lindl.) Bruttini 1930
= robur var. szovitzii A.DC 1864
= x szovitzii (A.DC) Zabel 1903, nom. illeg.
= x atropatena O.Schwarz 1938
= x komarovii A.Camus 1939
shrub (2-5 m ) ; crown fastigiate;
leaves : 6-9 cm long, 3-4cm wide; thick, oval, narrow, leathery, glabrous on both sides; margin with 5-9 pairs of shallow and narrow lobes; base weakly cordate or truncate;
fruit : cylindrical acorn, 2-3 cm long; flat cupule 2cm in diameter; maturing in 1 year;
grey bark remotely furrowed; hardy; good on silicious soils;
East Turquey to North Iran;
n° 158 in A.Camus;
x mazei macrocarpa X gambelii
= "Colorado foothills oak"
Colorado (native hybrid)

x megaleia Laughlin1962 lyrata x macrocarpa
Missouri ;
x mellichampii Trel. ex Sarg. 1918 laevis x laurifolia (or hemisphaerica ?)
= laurifolia var. tridentata Sarg. 1918
South Carolina, Florida ;
x mongolicodentata Nakai 1926

dentata x mongolica
= x kokaiensis A.Camus 1939
= x dentatomongolica Nakai 1926
China; Manshuria; Korea;


x moreha Kellogg 1863
(pro sp.)

also "x morehus"
kelloggii x wislizeni var. frutescens
= Q. x moreha Trel. 1917, nom. illeg.
"Oracle oak"
resembles Q. kelloggii, but its winter habit is recognizable by its evergreen (although few) leaves; dense foliage, lustrous green in Spring; 4-15 m tall; persistent ; leaves 4-12 cm with pointed lobes, glabrous above and minutely pubescent beneath; cupule as wislizeni but acorn broader; rare ; California ;

x morisii Borzi 1880 ilex x suber
= x bertrandii Albert & Reynier 1902
= x mixta Villalobos ex Colmeiro 1888
= x morisii f. mixta (Villar ex Colm.) Franco & Vasc. 1954
= x pseudomorisii A.Camus 1935
bark sometimes corky; twig whitish tomentose; bud small, tomentose, with persistent bracts;
leaves 3-7.5 x 2-4.5 cm, leathery, lustrous light green above, whitish tomentose beneath; apex pointed, base truncate or subcordate; margin dentate-serrate, with mucronate regular teeth, sometimes entire at basal half; few, prominent secondary veins; petiole tomentose 0.5-1.2 cm ; male flowers with longer, narrower stigmas than in Q. ilex; acorn with stiff, tomentose,1-3 cm long stalk; cup bell-shaped enclosing 1/2 of acorn, covered with numerous, silky scales with straight, free tip; maturing in 1 year;
France, Spain, Portugal, Italy, Sardinia, Sicily, Algeria;
-- For some Authors, Q. x mixta is more precisely the hybrid between Q. suber and Q. rotundifolia;
x moultonensis Ashe 1922 phellos x shumardii
= x shirlingii Bush ex Palm. & Steyerm.
= x hybrida Houba 1887
Alabama, Arkansas, Virginia, Tennessee ;
x munzii Tucker 1968 lobata x cornelius-mulleri
x mutabilis Palm. & Steyerm. 1935. palustris x shumardii
x neomairei A.Camus 1937 faginea x pyrenaica
= lusitanica f. ellipticifolia Cout. 1888
= alpestris var. ellipticifolia (Cout.) A.Camus 1939
= x mairei A.Camus 1935, nom. illeg.
= x numantina Ceballos & Vicioso 1942
= x welwitschii Sampaio 1947
shrub; bark greyish brown; young branchlet densely tomentose; bud 2-4 mm, with pinkish scales becoming glabrous; leaves 4.5-7.5 x 1.5-3.5 cm, almost evergreen, leathery, lustrous green above with a few stellate hairs, yellowish grey tomentose beneath; base rounded ou slightly cordate; margin with both lobes and sinuses narrow; 5-7 parallel vein pairs; petiole tomentose, pinkish, 4-15 mm long; fruit stalkless; cup half-round with numerous, appressed, densely yellowish tomentose scales;
northern Spain; Portugal ; Morocco ;
x neopalmeri Sudworth ex E.J.Palmer 1948 nigra x shumardii
S.E. USA ;
x neotharpii A.Camus 1939 minima x stellata
Texas ;
x nessiana E.J.Palmer bicolor x virginiana
artificial hybrid ( = Q. 'Nessiana') ;
x nipponica Koidz. 1912 dentata x serrata
= x angutelepidota Nakai 1917
= x yokohamensis Makino 1929
= x dentatoserratoides T.B.Lee 1961
Japan ; Korea ;
x organensis Trel. 1917 arizonica x grisea
from S.E. Arizona to W. Texas ;
x oviedoensis Sarg.1918 incana x myrtifolia
Florida ;
x palaeolithicola Trel. 1917 ellipsoidalis x velutina
Wisconsin, Indiana, Iowa ;
x palmeriana A.Camus 1953 falcata x imbricaria
= x anceps Palmer 1948, nom. illeg.
Eastern USA;
x parkeri A.Camus 1935 floribunda x leucotrichophora
North Pakistan ;
x pastorensis C.H.Muller 1936 clivicola x glaucoides
Mexico (Nuevo Leon) ;
x paui Vicioso 1950 pyrenaica x cerrioides
or fontqueri x cerrioides
dark pinkish twigs, more or less densely pubescent;
For Govaerts & Frodin, Q. x paui is the hybrid between Q. canariensis, Q. pyrenaica and Q. pubescens
North Spain;
x pendulina Kit. en Shulter 1814 robur x virgiliana
x podophylla Trel. 1917 incana x velutina
North Carolina;
x pongtungensis Uyeki 1932 aliena x mongolica
= x maccormickomongolica T.B.Lee 1961
= x alienocrispula H.0hba 1989
x pseudinfectoria A.Camus 1935 infectoria x pubescens ssp crispata
x rehderi Trel. 1917 ilicifolia x velutina
x richteri Baenitz

rubra x palustris
leaves glabrous, as in rubra, but narrower and deeper incised; hardy; prefers lime-free soils; handsome sturdy tree; fertile seeds;

x riparia Laughlin 1963 rubra x shumardii
Missouri ;
x robbinsii Trel. 1917 coccinea x ilicifolia
x robusta C.H.Muller 1934

emoryi x gravesii    (A. Camus : 428)
= Chisos mountains oak
Texas (Chisos Mountains, in wet canyons), to 1500 m ; reaches 13 m ; bark dark and furrowed; twigs dark brown, slender; leaves oval, 6-12 cm long, 2-5cm wide, apex pointed, base subcordate, toothed or rarely entire, glabrous on both sides; 8 vein pairs; petiole 1-2cm long, pubescent; acorn maturing in 2 years, 1-2cm long, enclosed for 1/4 or 1/3 of length in cup with pointed, appressed scales;
For C.H. Muller, and for Govaerts & Frodin (1998), it's a true species (fixed hybrid);

x rolfsii Small1905
(pro sp.)
chapmanii x minima
= geminata f. rolfsii (Small) Trel. 1925
rhizomatous bush with leaves intermediate between parents; acorn resembling the one of Q.minima; cotyledons connate (as for minima), while those of chapmanii are free;
S.E. USA ;
x rosacea Bechstein 1813 petraea x robur
= pedunculata var. rosacea (Bechst.)
Bluff & Fingerh. 1825
= x allardii Hy 1895
= x bossebovii Hy 1895
= x brevipes Heuffel 1850
= x hybrida Bechstein 1816
= x feketei Simonk. 1887
= x intermedia Boenningh. in Reichenb. 1831
= x jahnii Simonk. 1890
= x secalliana C.Vicioso 1950
= x sessiloïdes Léveillé 1917
Native hybrid in the whole Europe; hardy; all types of soils; characters intermediate between parents;
leaves are variable in size and shape; lobes are more regular than in Q. petraea; petiole glabrous,
0.5-2.5 cm long; male catkins with slightly pubescent axis; acorn's peduncle is the same length as the petiole of the nearest leave; convex scales cup; sometimes exist, on the same tree, petioles as in Q. petraea and peduncles as in Q.robur; hardy ; all types of soils;

-- var. filicifolia
= robur 'filicifolia'
with leaves divided into narrow segments, looking like penniform; dawny beneath as in Q. petraea; petiole elongated; stalked acorn cup;

-- Q. secalliana C.Vicioso 1950
= Q. x rosacea ssp secalliana (C.Vicioso) Rivas-Mart. & Saenz
 is a hybrid between Q. petraea and subspecies broteroana of Q. robur.
leaves 8-13 x 3.5-5 cm, both sides and midrib hairless; sinusal veins not only at the basal part; petiole hairless; acorn subglobose, almost all enclosed by cup;
x rotensis Penas, Llamas, Pérez Morales & Acedo 1997

pauciradiata X pyrenaica


x rudkinii Britt. 1882

marilandica x phellos
= dubia Ashe 1894, non Trel.
= phellos var. subimbricaria A.DC 1864
= x subimbricaria (A.DC) Sudw. 1897
Eastern USA to Texas ;

x runcinata (A.DC) Engelm. 1877

imbricaria x rubra
= rubra var. runcinata A.DC
Central and N.E. USA ;