Oaks of the World

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 Quercus spinosa
AuthorDavid ex Franch. 1884
Synonymsbullata Seemen 1897
ilex var. bullata (Seemen) Franch. 1899
gilliana Rehd. & Wils. 1916 (Camus : n 98)
ilex L. var. spinosa (David ) Franchet 1899
ilicioides David 1898
semecarpifolia var. spinosa (David ) Schottky 1912
Local namesci ye gao shan li ;
Range China; Myanmar; Taiwan; 1000-3100 m; introduced in Europe by Wilson (under the name Q.gilliana);
Growth habit4-10 metres tall (may reach 20 m);
Leaves 2.5-7 x 1.5-4 cm; lasting 2 years; oboval to elliptic; leathery; dark green; apex obtuse (sometimes pointed in Taiwan, see below); base subcordate; bullate; margins toothed or sometimes entire; base of midrib densely hairy (simple and fascicled hairs) beneath; adaxially some multiradiate hairs; 4-8 vein pairs, sunken below; petiole 1-3 mm long;
FlowersMay to August; pistillate inflorescences 1-3 cm long;
Fruitsacorn 1.2-2 cm long, 1 cm wide; ovoid globose; enclosed 1/3 by cup; cup with triangular, 1-1.5 mm long, grey pubescent (except near apex) scales ; maturing in 2 years;

Bark, twigs and
buds

young twigs covered with yellow brown stellate hairs, glabrescent;
Hardiness zone, habitathardy; mountain forests;
Miscellaneous -- A.Camus n 97;
-- Belongs to the Ilex Group, according to Denk & Grimm 2010;
-- For Govaerts & Frodin, 1998, Q. gilliana is a true species, whose leaves lack hairs at base of midrib beneath ; Tibet and in China (Sichuan, Yunnan, Gansu).
-- See the Key of Asian similar Quercus.
Subspecies and
varieties
There are 2 subspecies :
1/ subsp spinosa
the type described

2/ subsp miyabei (Hayata) A.Camus 1938
= Q.miyabei Kawak. ex Hayata 1918
= Q. spinosa var. miyabei Hayata 1918
= Q. semecarpifolia var. miyabei (Hayata) S.S.Ying 1988
= Q.taiyunensis Ling
= Q.tatakaensis Tomiya 1944
= Q. semecarpifolia f. tatakaensis (Tomiya) J.C.Liao 1970
named after plants from Taiwan Mountains; larger leaves (6.5-15 cm) with pointed apex; as these plants occur among populations of the type species, their taxonomic rank is uncertain...
Pictures

drawing 1
drawing 2